You can't clean a paper air filter with compressed air, it doesn't work like that unfortunately.
I was gonna say you might have an air leak but if you have a blocked/clogged up air filter then that will cause issues too.
And you can't tell if it's blocked by looking at it either, unless it really is filthy.
With all the work you've done i'm surprised you haven't changed it anyway just to eliminate it.
The thing is, if you do a carb balance with a clogged up air filter then you get a false reading, so the carbs will be adjusted wrongly making the bike run badly when you get it on the road. I've done it myself before but with a cheap air filter that wasn't letting enough air through, i changed it a few days after fitting and had to do the balance again, then everything was fine.
Although i did replace it with a K&N which does have much more airflow (and you can clean them with detergent and water).
I'm not saying this is your problem, but it is something that i would certainly consider changing, just to tick it off the list, plus if you don't know the history of the air filter then it won't hurt anyway.