Shouldn't make no difference as the bearings are roller bearings.
Does not matter as they are tightened onto the collar/spacer not the needle roller bearings
it is nothing to with having a collar/spacer as bushes also have a spacer, a bearing can be tightened in any position but a rubber bush should be tightened in its resting position under load otherwise the rubber bonding will get damaged when moved to its working position, not that will affect many motorcycles.